Five people accused of being connected to a £61m cannabis ring have been described in court as the lorry driver, girlfriend, the major dealer, minor dealer and courier.

All five deny being connected to a network of bosses smuggling £850,000 of cannabis a week from Holland, hidden in flower boxes, and storing drugs and money in lock-ups and homes in Kingston, Sutton, Ashtead and Epsom.

During his summing up at Southwark Crown Court on Wednesday prosecutor Timothy Cray QC asked the jury to decide whether they had been duped into their participation.

Was Lindsay Graham, 42, of Bushey Road, Sutton, the "major drug-dealer" who sold £2m worth of skunk cannabis and was referred to in deal books as Gyp?

And they were asked, was he in on a delivery to his Mitcham lock-up from convicted drug-dealers Peter Moran and Mark Kinnimont?

Challenging Mr Graham’s claim he was duped, Mr Cray said: "He tells you ‘well I used to trust people and take them at face value but now all that’s changed’ - a trusting, open approach to the world not traditionally associated with second-hand car dealers and rent collecters."

And questioning whether Graham was Gyp, he said Graham was "holding all the Aces" because he knew all the bosses of the network.

"Minor drug-dealer" James Hay, 31, of Stonny Croft, Ashtead, admits buying drugs at the Queen's Head pub in Epsom but claimed the dealer must have been spooked by undercover police into dumping 20kg of drugs into the boot of Mr Hay’s car.

Mr Hay said he was "stitched up" but in summing up Mr Cray said: "He is a self-confessed drug dealer in the past and admits he is an extremely experienced drug buyer.

"If he is telling the truth wouldn’t he of all people have realised something was up and he was being watched and he would have had nothing to do with it?"

Mandy Cripps, 34, the girlfriend of High Wycombe drug-dealer Robert Alexander, admitted recording details of the drug deals in a notebook found in her handbag, but claimed that her dyslexic boyfriend had dictated the figures to her.

Mr Cray said: "She asks you to set aside the natural inference on the basis that she was never more than a human ballpoint pen and gave no thought to what she was writing."

Of "driver" Richard Seville, 42, of Ecclesbourne Road, North London, the court heard claims he must have known the 27 deliveries to a lay-by near the A217 were nothing to do with the flower trade.

Mr Cray said: "He must have passed on enough of the yellow leaf flowers to cover Sherwood Forest.

"Did he know it was the drugs trade or did it pass him by?"

Discussing "courier" Peter Gilmour, 36, of High Wycombe, Mr Cray said: "Loyalty can be bought.

"Someone like Peter Gilmour can, with regret no doubt, put aside his principles."

The trial continues.